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You can use your first workflow to perform the manipulation of the values needed for the drop down which then triggers the second workflow. This will prevent any conflicts that may occur by actually using tools between the input file and the interface tool which is not recommended.
Using a chained app would definitely resolve the issue because essentially you are just doing a run in the background to build out the drop downs without the underscores. The user wouldnt really know what is happening.
The only other option would be to remove the underscores at the source.